Stuttgarter Beiträge zur Naturkunde Serie A (Biologie)

Herausgeber:

Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart

Stuttgarter Beitr. Naturk. Nr. 700 | 64S., 146 Abb. | Stuttgart, 10. TV. 2007

New species and additional records of Staphylinidae from Turkey V (Coleoptera)

VOLKER ASSING

Abstract

14 species of Staphylinidae from Turkey are described and illustrated: Aploderus lydicus n.sp. (Aydin, Izmir), Astenus (Eurysunius) occiduus n.sp. (Denizli, Aydin, Izmir), Medon reliquus n.sp. (Aydin, Izmir), Quedius (Raphirus) harpago n.sp. (Izmir), Cypha spathula n.sp. (Turkey: Manisa, Mersin, Osmanıye, Kahramanmaras, Antakya; Spain; Portugal), My/- laena nemorivaga n. sp. (Rize), Atheta (Datomicra) dissimulans n.sp. (Gümüshane), Drusilla lydica n.sp. (Aydin, Izmir), Pella gibbera n.sp. (Izmir), Pyroglossa pontica n.sp. (Rize), Cousya schuelkei n. sp. (Rize), Oxypoda (Oxypoda) subspectabilis n. sp. (Ordu), Dinusa smyr- nensis n.sp. (Izmir), and Pseudocalea messorphila n.sp. (Izmir). Gymnusa anatolica Korge, 1971, originally described as a subspecies of G. variegata Kiesenwetter, 1845, is regarded as a distinct species. Megalogastria Bernhauer, 1901, previously a subgenus of Aleochara Graven- horst, 1802, is elevated to genus; both the genus and its type species Aleochara cingulata Ep- pelsheim, 1889 are redescribed. A lectotype is designated for Aleochara cingulata. The male genitalia of Xantholinus osellai Bordoni (holotype), Gymnusa anatolica, and Atheta epirotica Benick are illustrated. Numerous additional records of previously described species are re- ported from Turkey, some also from Krasnodar (Russia), among them 74 first records from Turkey, one from Anatolia, one from Russia, and three from the Russian South European ter- ritory. The distributions of 14 species are mapped.

Keywords: Coleoptera, Staphylinidae, taxonomy, new species, new records, lectotype de- signation, Turkey, Russia, Spain, Portugal.

Zusammenfassung

14 Arten der Familie Staphylinidae aus der Türkei werden beschrieben und abgebildet: Aploderus lydicus n.sp. (Aydin, Izmir), Astenus (Eurysunius) occiduus n.sp. (Denizli, Aydin, Izmir), Medon reliquus n. sp. (Aydin, Izmir), Quedius (Raphirus) harpago n.sp. (Izmir), Cy- pha spathula n. sp. (Türkei: Manisa, Mersin, Osmaniye, Kahramanmaras, Antakya; Spanien; Portugal), Myllaena nemorivaga n.sp. (Rize), Atheta (Datomicra) dissimulans n.sp. (Gümüshane), Drusilla lydica n.sp. (Aydin, Izmir), Pella gibbera n.sp. (Izmir), Pyroglossa pontica n. sp. (Rize), Cousya schuelkei n. sp. (Rize), Oxypoda (Oxypoda) subspectabilis n. sp. (Ordu), Dinusa smyrnensis n. sp. (Izmir) und Pseudocalea messorphila n. sp. (Izmir). Gymnu- sa anatolica Korge, 1971, urspriinglich als Unterart von G. variegata Kiesenwetter, 1845 be- schrieben, wird als distinkte Art betrachtet. Megalogastria Bernhauer, 1901, bisher Untergat- tung von Aleochara Gravenhorst, 1802, wird zur Gattung erhoben und einschließlich ihrer Typusart Aleochara cingulata Eppelsheim, 1889 redeskribiert. Für Aleochara cingulata wird ein Lectotypus designiert. Die männlichen Genitalien von Xantholinus osellai Bordoni (Ho-

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lotypus), Gymnusa anatolica und Atheta epirotica Benick werden abgebildet. Zahlreiche wei- tere beschriebene Arten werden aus der Türkei, einige auch aus Krasnodar (Russland) gemel- det, darunter 74 Erstnachweise für die Türkei, einer für Anatolien, einer für Russland und drei für das südeuropäische Russland (Russian South European territory). Die derzeit bekannte Verbreitung von 14 Arten wird anhand von Karten illustriert.

Contents 199] Fats gore heres sta) re Te On ee a en Ed 2 2= Material, merbhads Sand abbreviatisns et a a ER En en er a T 3 3. Descriptions of new species and. additional records». ...2.2.2... Fide ee ec een 3 DR eterence Ss a LE Een U Le a, «Nat Oe nd en 61

1 Introduction

As has been discussed in more detail earlier, the highly diverse Staphylinidae fau- na of Turkey had for a long time been neglected, but has received more attention recently (e.g. AssING 2003a, 2004a, 2004b, 2006f). Approximately 1400 species are currently known from Turkish territory; almost 200 of them were described only in the past decade (e.g. AssING 2003a, 2004a, 2004b, 2006f, 2006h, 20061, SMETANA 2004). Disregarding the Pselaphinae, the staphylinid genera with the highest diver- sity of endemic species in Turkey are Geostiba Thomson (63 species), Leptusa Kraatz (28 species and subspecies), and Sunius Curtis (25 species) (AssING 2003b, 2004e, 2006f, 2006h, 20061, in press). Unsurprisingly, numerous recently described taxa refer to these genera. However, a large number of new species have also been described in various other genera. At present, the rate of additional discoveries is still increasing rather than decreasing. The same applies to records of described - and often widespread species which were previously unknown from Turkish ter- ritory.

The present paper is based primarily on material collected during six field trips to Turkey carried out by ALEXEY SOLODOVNIKOv (currently Chicago) in June 1998, by the author in December 2005, by HEINRICH MEYBOHM (Stelle) in March 2006, by PauL WUNDERLE (Mönchengladbach) and the author in April 2006, by VOLKER BRACHAT (Geretsried) and HEINRICH MEYBOHM in April 2006, as well as by MICHAEL SCHULKE (Berlin) and the author in July/August 2006. This article is the fifth part of a series dealing with Turkish records of Staphylinidae of various genera and subfamilies. The new species and records of Leptobium Casey, Sunius, Lathro- bium Gravenhorst, Leptusa, Geostiba, and some species of Oxypoda Mannerheim have been - or will be treated separately elsewhere (AssınG 2006a, 2006d, 2006h, 2006i, in press, in prep.). Practically all the Tachyporinae and some Oxytelinae will be studied by SCHULKE (in prep.). The present paper provides descriptions of 14 new species belonging to 14 different genera of the Oxytelinae (one species), the Paederi- nae (two species), the Staphylininae (one species), and especially of the Aleocharinae (10 species). In addition, numerous records of zoogeographic interest are compiled, among them as many as 74 first records of previously described species from Turk- ish territory.

Acknowledgements

My sincere thanks are due to VOLKER BRACHAT, HEINRICH MEYBOHM, MICHAEL SCHÜLKE, ALEXEY SOLODOVNIKOV, and PAUL WUNDERLE for the generous gift and loan, re-

ASSING, STAPHYLINIDAE FROM TURKEY V 3

spectively, of material for the present study. JÜRGEN VOGEL (Görlitz) assisted with the iden- tification of some Atheta. I am also indepted to JOHANNES REIBNITZz (Stuttgart) for improv- ing the quality of some of the figures.

2 Material, methods, and abbreviations

The material treated in the present paper is deposited in the following public institutions and private collections:

cApf private collection W. Apreı, Eisenach

cAss private collection V. Asstnc, Hannover

cFel private collection B. FELDMANN, Münster

cSch private collection M. SCHÜLkE, Berlin

cSol private collection A. SOLODOVNIKOv, Chicago

cWun private collection P. WUNDERLE, Mönchengladbach

DEI Deutsches Entomologisches Institut, Müncheberg (L. ZERCHE)

MCSNV Museo Civico di Storia Naturale Verona (L. LATELLA) NHMW Naturhistorisches Museum, Wien (H. SCHILLHAMMER) NMP Narodni Muzeum v Praze (J. Hayex)

The morphological studies and drawings were carried out using a Stemi SV 11 microscope (Zeiss Germany) and a Jenalab compound microscope (Carl Zeiss Jena) with a drawing tube. For the photographs a digital camera (Nikon Coolpix 995) was used. The maps were generat- ed using the online generic mapping tool (GMT) of the Geomar website at www.aquarius. geomar.de/omc.

The following morphological parameters were examined:

AL length of antenna

AW maximal width of abdomen

EL length of elytra along suture, from apex of scutellum to posterior margin EW combined width of elytra

HL head length from anterior margin of clypeus to neck

HW maximal head width

ML length of (median lobe of) aedeagus from base to apex of ventral process BL length of pronotum along midline

PW maximal width of pronotum

Tal length of metatarsus

Tale length of metatibia

AuB body length from mandibles to apex of abdomen

The genera are arranged by subfamily; the order of subfamilies is in accordance with the checklist of Central European Staphylinidae in AssING & SCHULKE (2001). Within genera, the species are grouped according to their current subgeneric assignment or their affiliations to certain species groups.

In the material sections, the Turkish provinces are arranged as follows: 1. provinces of northern Turkey from west to east; 2. provinces of western, southwestern, and southern Ana- tolia from west to east.

Labels of type material are cited in their original spelling and language, except for the fol- lowing minor adaptations according to the general requirements of the journal: names of col- lectors in small capitals, scientific names of genera and species in italics, dates with the months always in Roman numbers.

All the material listed in chapter 3 is from Turkey, unless stated otherwise.

3 Descriptions of new species and additional records

Micropeplus fulvus Erichson, 1840

Material examined: Manisa: 1 ex., Boz Daglar, SE Turgutlu, 38°24N, 27°52E, 800 m, ceme- tery, litter of oak and shrubs, 24.X11.2005, leg. AsstnG (cAss); 1 ex., Sipil Dag Milli Park,

4 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

38°33N, 27°28E, 1080 m, 25.1V.2006, leg. BRACHAT & MEYBOHM (cAss). Izmir: 1 ex., Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°06E, 1480m, N-slope with grass and stones, sifted, 3.IV.2006, leg. AssınG (cAss); 1 ex., Bozdag, 38°20N, 28°06E, 1300-1560 m, 21.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Nif Dagi, 38°24N, 27°24E, 1010 m, 23.1V.2006, leg. BRACHAT & MEYBOHM (cAss). Aydin: 4 exs., ca. 110km S Izmir, WSW Soke, Dilek Dagi, 37°39'23N, 27°08’14E, 950 m, N-slope, pine and oak litter, grass roots sift- ed, 25.X11.2005, leg. Asstnc (cAss); 1 ex., Dilek Dagi, Kanyon, 37°41N, 27°10E, 70-370 m, 16.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 2 exs., Dilek Dag, S Kanyon, 37°40N, 27°11E, 670m, 17.IV.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., ca. 20km NE Kuyucak, Bayrak Tepe, 37°58N, 28°34E, 900m, N-slope, oak litter and grass sifted, 7.TV.2006, leg. AssınG (cAss); 1 ex., NE Aydin, Imambaba Tepesi, 37°57N, 27°54E, 1460m, 20.IV.2006, leg. BRACHAT & MEYBOHM (cAss). Mugla: 8 exs., 16km N Milas, Labranda, 37°25N, 27°43E, 670m, 21.11.2006, leg. MEYBOHM (cAss); 4 exs., Labranda, 37°25N, 27°49E, 550-660 m, 18.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., S Bafa lake, 37°28N, 27°24E, 170m, 26.11.2006, leg. MEYBOHM (cAss); 1 ex., Yatagan-Bozdogan, 37°27N, 28°18E, 825m, 19.IV.2006, leg. BRACHAT & MEYBOHM (cAss); 3 exs., Yatagan-Bozdogan, 37°38N, 28°19E, 590 m, 19.TV.2006, leg. BRACHAT & MEYBOHM (cAss); 7 exs., ca. 20 km NNE Fethiye, 36°47N, 29°11E, 970m, mixed forest, 27.III.2002, leg. AssınG (cAss); 2 exs., SE Fethiye, Baba Dag, above Ovacik, 36°33N, 29°11E, 1170 m, pine, cedar, and Quercus ilex litter, 30.111.2002, leg. AssING (cAss); 2 exs., SE Fethiye, Baba Dag, above Ovacik, 36°33N, 29°10E, 680 m, pine lit- ter, 30.111.2002, leg. AssınG (cAss); 9 exs., W Kemer, road to Ovacik, 36°36N, 30°29E, 325 m, litter of oak and other deciduous trees, 2. IV.2002, leg. AssınG (cAss). - Antalya: trex." Kalkan, Dumanlı Dagi, 36°24N, 29°26E, 1230m, forest margin, grass and moss sifted, 5.X.2002, leg. AssınG (cAss); 198 exs., ca. 20km N Kas, S Karaovabeli Gec., 36°23N, 29°43E, 830 m, paved oak and pine forest, sifted: 26.111.2002, leg. Assıng (cAss). Mersin: 1 ex., 20 ian NNW Mut, 36°49N, 33°19E, 1320 m, pine forest with Quercus ilex, 25.X11.2000, leg. AssınG (cAss); 28 exs., road from Silifke to Gülnar, ca. 40 km W Silifke, 36°21N, 33°35E, 1015 m, oak litter sifted, 27.XII.2000, leg. AsstnG (cAss); 1 ex., road Silifke-Gülnar, 36°21N, 33°35E, 1000 m, 6.V.2004, leg. BRACHAT & MFEYBOHM (cAss); 1 ex., road to Güzeloluk, S Aydınlar, 36°42N, 34°10E, 1110 m, 5.V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., 23km N Silifke, 36°32N, 33°56E, 970 m, 18.1V.2005, leg. BRACHAT & MEYBOHM (cAss); 3 exs., NW Silifke, road Mut-Ermenek, 15km before Ermenek, 36°38N, 33°01E, 1030 m, 20.IV.2005, leg. BRACHAT & MEYBOHM (cAss); 1 ex., NW Tarsus, road Camlıyayla-Gözne, 37°06N, 34°37E, 570-610 m, 25.1V.2005, leg. BRACHAT & MEYBOHM (cAss). Osmaniye: 1 ex., 15km E Os- maniye, NW Yarpuz, 37°04N, 36°26E, 920 m, stream bank, alder litter, 11.IV.2002, leg. AssınG (cAss); 1 ex., NE Kadirli, Torlar near Andirin, 37°33N, 36°26E, 1110 m, 30.IV.2005, leg. BRACHAT & MEYBOHM (cAss). Antakya: 1 ex., 9km SE Iskenderun, 6km NE Belen, 36°32N, 36°15E, 1480 m, edge of snowfield, under stones and sifted, 4.TV.2004, leg. AssınG (cAss); 1 ex., 9km SE Iskenderun, 5km NE Belen, 36°31N, 36°15E, 1240 m, mixed oak and beech forest, sifted, 4.IV.2004, leg. Asstnc (cAss); 1 ex., Ziyaret Dagi, W Senköy, 36°01N, 36°07E, 750 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Ziyaret Dagi, W Sungur, 36°00N, 36°06E, 660 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 6 exs., Ziyaret Dagı, W Sungur, 35°59N, 36°05E, 710 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Zi- yaret Dagi, Leylekli, 35°58N, 36°03E, 510 m, 22.1V.2004, leg. BRACHAT & MEYBOHM (cAss). Kahramanmaras: 1 ex., 20km SW Hopurlu, 520 m, 37°28’45N, 36°48’10E, 27.IV.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., 50km W Kahramanmaras, Baskonus Yaylası, 37°34N, 36°34E, 1250 m, 5.V.2005, leg. BRACHAT & MEYBOHM (cAss). Gaziantep: 1 ex., Kartal Dagı, W Yamacoba, 37°10N, 37°05E, 1200 m, 25.IV.2004, leg. BRACHAT & MEYBOHM (cAss).

According to SMETANA (2004), this common species was previously unknown from Turkey.

Micropeplus tesserula Curtis, 1828

Material examined: Mersin: 1 ex., road from Silifke to Gülnar, ca. 40km W Silifke, 36°21N, 33°35E, 1015 m, oak litter sifted, 27.XII.2000, leg. AsstNc (cAss).

ASSING, STAPHYLINIDAE FROM TURKEY V 5

Micropeplus tesserula is widely distributed in the Palaearctic region, but had not been reported from Turkey (HERMAN 2001, SMETANA 2004).

Proteinus meuseli Dauphin, 1995

Material examined: Izmir: 1 ex., Bozdag, 38°20N, 28°06E, 1300-1560 m, 21.IV.2006, leg. BRACHAT & MEYBOHM (cAss).

The species was previously known only from Croatia, Romania, Bosnia-Herze- govina, and Greece (Assinc 2004d, DaupHIN 1995). It is here reported from Turkey for the first time.

Philorinum sordidum (Stephens, 1834)

Material examined: Manisa: 1 ex., Sipil Dagi Milli Parki, 38°33N, 27°28E, 1080m, 25.1V.2006, leg. BRACHAT & MEYBOHM (cAss).

According to SMETANA (2004), this common species was previously unknown from Turkey.

Omalium assingi Zanetti, 2002

Material examined: Manisa: 1 ex., Boz Daglar, SE Turgutlu, 38°24N, 27°52E, 800 m, ceme- tery, litter of oak and shrubs, 24.X1I.2005, leg. AssınG (cAss).

This species has become known only from southern and western Anatolia, where it is apparently widespread and not uncommon, especially at higher altitudes. A dis- tribution map is provided by Assinc (2004b).

Omalium cribriceps Fauvel, 1900

Material examined: Izmir: 1 ex., N Izmir, Yamanlar Dagi, 38°33N, 27°09E, 940 m, grassy patch in pine forest, under stones, sifted grass roots, 28.XII.2005, leg. AssınG (cAss); 1 ex., Nif Dagi, 38°24N, 27°24E, 1010m, 23.1V.2006, leg. BRACHAT & MEYBOHM (cAss). Mugla: 4 exs., ca. 20km SW Mugla, N Meke, 37°13N, 28°12E, 590 m, pasture with stones at roadside, sifted and under stones, 12.IV.2006, leg. Assıng, WUNDERLE (cAss, cWun); 1 ex., ca. 25 km SW Mugla, 37°11N, 28°06E, 1130 m, N-slope, pine forest, grass roots and pine litter sifted, under stones, 12.IV.2006, leg. WUNDERLE (cWun); 2 exs., Yatagan-Bozdogan, 37°26'35N, 28°18’07E, 825 m, 19.IV.2006, leg. BRACHAT & MEYBOHM (cAss). Ankara: 4 exs., Gölbası, 750 m, 29.X.1995, leg. Vir (cAss).

In Turkey, O. cribriceps was previously known only from the Van lake area and from western Antalya province (AssınG 2004b, ZANETTI 2002).

Mannerheimia brevipennis (Motschulsky, 1860)

Material examined: Ankara: 8 exs., SE Ankara, N-Elma Daßı, 1600 m, moss and litter sift- ed, 31.X.1995, leg. Vrr (cAss). - Gümüshane: 4 exs., ca. 25km SW Gümüshane, Tersundagi Gec., 40°18N, 39°18E, 2070m, N-slope, spruce forest, litter and dead wood sifted, 24.VII.2006, leg. AssınG (cAss).

In Turkey, this rare species is now known from Ankara, Kahramanmaras, Adıya- man, Gümüshane, and Kayseri provinces (AssING 2004b, 2006f).

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Anthobium fusculum (Erichson, 1839)

Material examined: Aydin: 1 ex., ca. 15 NNE km Aydin, Imambaba Tepesi, 37°57N, 27°54E, below peak 1600 m, edge of pine forest with grass, shrubs, sifted and under stones, 5.1V.2006, leg. AssınG (cAss). - Erzurum: 1 ex., 30-45km NNE Erzurum, Dumludagi, ca. 40°11N, 41°27E, 2500-2900 m, 15.V1.1998, leg. SOLODOVNIKOV (cAss).

In Turkey, the species was previously known only from Mersin, Adana, and Adıyaman provinces (AssınG 2004b, 2006f).

Acidota crenata (Fabricius, 1793)

Material examined: Rize: 5 exs., 40km SSW Hopa, source of Caglayan D., ca. 41°06N, 41°22E, 2700-2900 m, 25.V1.1998, leg. SOLODOVNIKOV (cAss).

Acidota crenata is widespread in the Palaearctic region, but was previously un- known from Turkish territory (SMETANA 2004).

Aploderus lydicus n.sp. (Figs. 1-12, 50)

Types Holotype 6: N37°56'47 E027°53'53 (8), Türkei, Aydin, Pasayaylasi, 1460 m, 20.1V.2006, l. BRACHAT & MEYBoHM / Holotypus 6 Aploderus lydicus sp. n. det. V. AsstNG 2006 (cAss). Paratypes: 16, 4 PP: same data as holotype (cAss); 1d: TR Prov.: Aydin (8), N Aydin, Pasayaylasi, 1460 m, 20.1V.2006, N37°56’47", E27°53'53", leg. MEYBOHM & BRACHAT (cAss); 1 2: N38°35’25 E026°29'02 (22), Türkei, Izmir, Karaburun, 550 m, 26.1V.2006, |. BRACHAT & MEYBOHM (cAss); 1 2: 28.XI1.2005, Yamanlar, 940 m, Izmir, S. ANLAs (cAss).

Etymology The name (adjective) is derived from Lydia, the ancient name of the region where the species was found.

Description

TL: 3.5-4.5mm. Habitus as in Fig.1. Coloration: head and abdomen black; pronotum dark brown with paler lateral and posterior margins; elytra brown to dark brown; legs yellowish; antennae dark brown.

Head with sexual dimorphism, larger in ¢ than in $ (Figs. 3-6); posterior sulcus absent; posterior and lateral areas with rather coarse puncturation of rather variable density, interstices on average approximately as wide as diameter of punctures; frons impunctate; integument shiny, usually with shallow microreticulation at least in pos- terior area, frons with or without transverse microstriae; eyes relatively small, ap- proximately as long as (2 and small 2) or shorter (large 3) than postocular region in dorsal view, and weakly prominent (Figs. 7-8). Antenna as in Fig. 2.

Pronotum distinctly transverse, 1.35 (small 2) to 1.45 (large d) times as wide as long and 1.05 (large 3) to 1.20 (@) times as wide as head (Figs. 3-6); posterior angles completely rounded, not marked; on either side of impunctate midline with ill- defined shallow oblong impressions with coarse and dense, sometimes ill-defined puncturation; in lateral areas with less dense puncturation; integument with shallow longitudinally striate microsculpture.

Elytra as wide (large d) or up to 1.1 times as wide (small 3) and at suture 1.15-1.20 times as long as pronotum (Figs. 3-6); puncturation dense and coarse, in-

ASSING, STAPHYLINIDAE FROM TURKEY V /

Figs. 1-12. Aploderus lydicus n. sp. 1. Male habitus (holotype). 2. Antenna. 3-6. Forebody of three males (3-5) and female (6). 7-8. Head in lateral view ot large male (7) and small male (8). 9. Male sternite VIII. 10-12. Aedeagus in lateral and in ventral view. Scale bars: 1 mm (1), 0.5mm (2-8), 0.2 mm (9), 0.1 mm (10-12).

terstices on average narrower than diameter of punctures and without microsculp- ture. Hind wings present, length not examined.

Abdomen as wide as or slightly wider than elytra, widest at segments V—-VI; punc- turation sparse and fine; tergites with distinct microreticulation; posterior margin of tergite VII broadly concave and with palisade fringe; posterior margin of tergite VIII weakly concave.

3: posterior margin of sternite VIII broadly concave (Fig.9); aedeagus as in Figs. 10-12.

8 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

Comparative notes

Four Aploderus species were previously known from the Western Palaearctic re- gion, two of them widespread: the Palaearctic A. caelatus (Gravenhorst), the Euro- pean A. caesus (Erichson), A. endogaeus Assing from southwestern Anatolia (Mugla, Antalya), and the microphthalmous A. schweigeri (Smetana) from northern Anato- lia (AssING 2003a, HERMAN 2001, MAKRANCzy 2006, SMETANA 1967). In addition to the morphology of the aedeagus, the new species is distinguished from these species by the following characters:

- from A. caelatus by the much smaller eyes, the more distinct and coarser punc- turation and the much shallower microsculpture of head and pronotum, as well as by the coarser puncturation of the distinctly shorter and narrower elytra;

- from A. caesus by smaller average size, shorter antennae, smaller and less promi- nent eyes; the absence of a posterior sulcus on the head, the coarser, but less well de- limited puncturation of head and pronotum, the much shorter pubescence of the forebody, the absence of distinct posterior pronotal angles (in A. caesus well- marked), as well as by the shorter and narrower elytra;

from A. endogaeus by larger size, darker coloration of pronotum, elytra, and ab- domen (in A. endogaeus yellowish brown to rufous), a larger and anteriorly more distinctly widened pronotum, and by broader and longer elytra;

- from A. schweigeri at once by the darker coloration, the completely different body shape (in A. schweigeri with abdomen distinctly enlarged), the distinctly larg- er eyes, more slender antennae (in A. schweigeri with distinctly transverse anten- nomeres VII-X), the more transverse pronotum, the much longer and broader ely- tra, and the presence of hind wings.

For illustrations of the habitus and the genitalia of A. schweigeri and A. endogaeus (male unknown) see SMETANA (1967) and AssING (2003a).

Intraspecific variation The species is subject to considerable intraspecific variation, especially in the males, affecting not only body size, but also the shape and the relative size of the head, the relative size of the eyes, the microsculpture and puncturation of the head, as well as the shape and relative size of the pronotum. For a comparison of three males of different sizes and a female see Figs. 3-8.

Distribution and bionomics The species was recorded in three localities in western Turkey (Aydin and Izmir provinces) (Fig.50). Apart from the altitude (550-1460 m), no bionomic data are available.

Carpelimus subtilis (Erichson, 1839) Material examined: Ankara: 2 exs., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss).

According to SMETANA (2004), this species had not been reported from Turkey before.

ASSING, STAPHYLINIDAE FROM TURKEY V 9

Oxytelus laqueatus (Marsham, 1802) Material examined: Rize: 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss).

Although widely distributed in the Palaearctic region and in the Americas, this species had been unknown from Turkey (HERMAN 2001).

Bledius subterraneus Erichson, 1839

Material examined: Trabzon: 32 exs., ca. 50 km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1540 m, bank of stream, 26.VII.2006, leg. Assıng, SCHULKE (cAss, cSch).

According to HERMAN (2001) and SMETANA (2004), this widespread Palaearctic species had not been reported from Turkey.

Stenus distortus Assing, 2006

Material examined: Gümüshane: 10 exs., ca. 50km SW Trabzon, 9-10 km S Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). Trabzon: 2 exs., ca. 50km S Of, S Uzungöl, 40°36N, 40°17E, 2050 m, 4.VIII.2006, leg. SCHULKE (cSch).

A map illustrating the distribution of this recently described species, which has become known only from eastern and central southern Anatolia, is provided by Ass- ING (2006b).

Stenus humilis Erichson, 1839 Material examined: Gümüshane: 2 exs., ca. 50km SW Trabzon, 9-10km S$ Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss, cSol).

This is the first record of S. humilis from Turkey.

Stenus trapezipennis Puthz, 1981

Material examined: Rize: 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, 28.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°15N, 41°13E, 500 m, 29.V1.1998, leg. SOLODOVNIKOV (cAss).

This species has been recorded only from northeastern Anatolia and Georgia (HERMAN 2001).

Rugilus subtilis (Erichson, 1840) Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss).

This is the first record of Rugilus subtilis trom Turkey.

Rugilus lesbius Assing, 2005

Material examined: Aydin: 2 exs., Dilek Dagi, Kanyon, 37°41N, 27°10E, 70-370 m, 16.1V.2006, leg. BRACHAT & MEYBOHM (cAss).

10 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

This recently described species had been known only from the Greek island Les- bos (AssING 2005d).

Astenus (Eurysunius) occiduus n.sp. (Figs. 13-21, 83)

Types

Holotype ¢ [with four workers of Tetramorium sp. attached to the same pin]: TR [21] - Denizli, 60 km E Mugla, S Kale, 1280 m, 37°25’39N, 28°53'18E, 11.IV.2006, V. Asstnc / Holo- typus d Astenus occiduus sp. n. det. V. AssınG 2006 (cAss).

Paratypes: 1d: same data as holotype (cAss); 14, 2 2?: TR [22] - Denizli, 60km E Mugla, S Kale, 1240 m, 37°23'12N, 28°53’41E, 11.IV.2006, V. AssınG (cAss); 1 2: TR [8] - Aydin, 15km NNE Aydin, Imambaba T., below peak, 1600m, 37°57'16N, 27°53'55E, 5.1V.2006, V. Asstnc (cAss); 1 2: TR [13] - Aydin, 20 km NE Kuyucak, Bayrak Tepe, 1480 m, 38°00’09N, 28°34'53E, 7.IV.2006, P. WUNDERLE (cWun); 12: TR Prov.: Izmir (12), NO Odemis, Boz Dag, 1560 m, 21.IV.2006, N38°20'11”, E28°6'26", leg. MEYBOHM & BRACHAT (cAss).

Etymology The name (Latin, adjective: western) alludes to the fact that this species is the westernmost representative of Eurysunius in Turkey.

Description

TL: 3.8-4.9 mm. Habitus as in Fig. 13. Coloration: forebody blackish, with the posterior 1/4-1/3 of elytra yellow, abdomen blackish with the narrow posterior mar- gins of the tergites and the apex somewhat paler, legs and antennae rufous.

Head transverse, 1.25-1.30 times as wide as long; dorsal surface distinctly convex, with very dense, large, but rather shallow punctures, and only with subdued shine (Fig. 14); pubescence short, greyish, and depressed; eyes relatively small and weakly prominent, temples approximately 1.7-2.5 times as long as eyes in dorsal view (Figs. 16-17). Antenna relatively stout, antennomeres V-X moderately oblong (Fig. 15).

Pronotum across anterior angles slightly (approximately 1.05 times) wider than head, 1.05-1.10 times as wide as long (width measured across anterior angles); max- imal width at anterior angles; surface without distinct impressions; posterior margin convex; lateral margins straight, each with two long setae of slightly more than half the length of lateral margin of pronotum (Fig. 14), one at anterior and one at poste- rior angle; microsculpture barely noticeable, almost absent; puncturation similar to that of head, but slightly sparser, surface somewhat more shiny than that of head; pubescence of similar length as that of head, but less fine and more conspicuous.

Elytra approximately 1.1 times as wide and at suture about 0.65 times as long as pronotum; microsculpture absent; puncturation very distinct and granulose; inter- stices approximately as wide as punctures; pubescence yellowish, more distinct than that of head and pronotum; long setae present only at posterior margin near poste- rior angles, absent from lateral margins. Hind wings reduced.

Abdomen about as wide as or slightly wider than elytra, widest at segments V-V]; puncturation distinct and granulose, somewhat denser on anterior than on posterior tergites; tergite VII with sparser puncturation, interstices on average twice as wide as punctures and without microsculpture; posterior margin of tergite VII with narrow rudiment of a palisade fringe.

ASSING, STAPHYLINIDAE FROM TURKEY V 11

20

Figs. 13-21. Astenus occiduus n. sp. 13. Habitus. 14. Forebody. 15. Antenna. 16-17. Head in lateral view of specimens from Denizli (16) and from Bozdag (17). 18-19. Aedeagus in lateral and in ventral view. 20-21. Apical part of aedeagus in ventral and in lateral view. - Scale bars: 1mm (13), 0.5mm (14-17), 0.2 mm (18-21).

3: sternite VII unmodified; posterior margin of sternite VIII with deep and acute incision; aedeagus as in Figs. 18-21.

Comparative notes According to a recent revision of the Eurysunius species of Turkey and the Cau- casus region, three representatives were known from Turkish territory, all of them from northern and eastern Anatolia (Kastamonu, Ordu, Kayseri) (AssING 2002c). From all these species, A. occiduus is distinguished by the morphology of the aedea- gus. In addition, it is separated from them as follows:

12 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 700

- from A. bicoloratus Assıng (Ordu) by the more slender antennae (with distinct- ly oblong antennomeres V—X, the straight lateral margins of the pronotum (in A. bi- coloratus sinuate), the presence of long setae at the posterior pronotal angles, and by the absence of lateral impressions on the pronotum;

- from A. paphlagonicus Assıng (Kastamonu) by less slender antennae, a trans- verse pronotum (in A. paphlagonicus about as wide as long) with sparser punctura- tion and straight lateral margins (in A. paphlagonicus weakly convex), and by the less extensive yellow coloration of the elytra (in A. paphlagonicus extending to the mid- dle of the elytra);

- from A. sexsetosus Assing (Kayseri) by less slender antennae, the absence of an additional long seta between the anterior and the posterior pronotal setae, the denser pronotal puncturation, as well as by the much less extensive yellow coloration of the elytra (in A. sexsetosus extending almost to the anterior margin).

For illustrations of the habitus and the genitalia of these species see AssING (2002c).

Remarks Whether or not the females from Aydin and Izmir are conspecific with the types from Denizli cannot be said with certainty. The eyes of these specimens are some- what smaller (Figs. 16-17), but since no additional significant differences were found, they are here considered an expression of intra- rather than interspecific vari- ation.

Distribution and bionomics

The Palaearctic subgenus Eurysunius currently includes some 45 species, most of which occur in the Western Mediterranean. Only 9 species had been recorded from the Eastern Mediterranean and the Caucasus region. Astenus occiduus is the fourth representative of the subgenus to become known from Turkey and the first Eury- sunius species from Western Turkey (Denizli, Aydin, Izmir) (Fig. 83). Records of species of this subgenus are generally rare; many taxa are still represented only by their respective holotypes. Recent observations suggest that all Eurysunius species may be associated with ants of the genus Tetramorium Mayr (AssING 2002c, 2003c). This hypothesis is here supported by new evidence. At least six of the type speci- mens were found in nests of a yellowish Tetramorium sp. All the records are from grassland biotopes at altitudes of 1240-1600 m.

Medon reliquus n.sp. (Figs. 22-29, 71)

Types

Holotype 6: TR [1] - Izmir, 1480m, Boz Daglar, above Bozdag, road to ski resort, 38°21'26N, 28°05’38E, 3.TV.2006, V. Asstnc / Holotypus d Medon reliquus sp. n. det. V. Ass- ING 2006 (cAss).

Paratypes:4 dd, 6 22: same data as holotype; 2 dd, 2 2: same data, but leg. WUNDER- LE (cWun); 1d, 1 2: TR [11] - Aydin, 20km NE Kuyucak, Bayrak Tepe, 850 m, 37°57'56N, 28°33'23E, 7.1V.2006, P. WUNDERLE (cWun); 1 2: TR [12] Aydin, 20km NE Kuyucak, Bayrak Tepe, 900 m, 37°58'06N, 28°33'33E, 7.1V.2006, P. WUNDERLE (cWun).

ASSING, STAPHYLINIDAE FROM TURKEY V 13

Figs. 22-29. Medon reliquus n. sp. 22. Habitus. 23. Forebody. 24. Antenna. 25. Head in lat- eral view. 26. Male sternite VII. 27. Male sternite VIII. 28-29. Aedeagus in lateral and in ven- tral view. Scale bars: 1 mm (22), 0.5 mm (23-25), 0.2 mm (26-27), 0.1 mm (28-29).

Etymology The name (Latin, adjective: remaining, relict) alludes to the fact that this addition to the Turkish Medon fauna was discovered only after the genus had been thoroughly revised.

Description

TL: 3.1-3.8 mm. Habitus as in Fig.22. Coloration: body uniformly rufous; legs and antennae pale reddish.

Head approximately as long as wide to weakly oblong; puncturation dense, coarse, and well-defined; interstices on average narrower than punctures and with- out microsculpture (Fig. 23); eyes rather small (Fig. 25) and not distinctly projecting from lateral outline of head, postocular region at least 3 times as long as eyes in dor- sal view. Antenna as in Fig. 24.

Pronotum approximately 1.1 times as wide as long and 1.1 times as wide as head; puncturation in lateral areas similar to that of head (Fig. 23).

Elytra approximately 1.1 times as wide and at suture 0.8 times as long as prono-

14 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

tum; puncturation ill-defined, finer and denser than that of head and pronotum (Fig. 23). Hind wings reduced.

Abdomen slightly wider than elytra, widest at segment VI; integument with dis- tinct microsculpture everywhere; puncturation fine and moderately dense; posterior margin of tergite VII without palisade fringe.

3: posterior margin of sternite VII almost truncate, with few modified dark setae in the middle, without palisade setae (Fig. 26); sternite VIII with relatively small pos- terior excision (Fig. 27); aedeagus as in Figs. 28-29.

Comparative notes and phylogenetics

The Medon species of the Western Palaearctic region were comprehensively re- vised only recently (AssING 2004c, 2006e). Medon reliquus combines an aedeagal morphology that is similar to that of some species of the M. apicalis group (M. ma- ronitus (Saulcy), M. beydaghensis Fagel) with a chaetotaxy of the male sternite VII that resembles that of species of the M. petrochilosi group (e.g. M. cerruti Coittait, M. impar Assing). This suggests that the M. apicalis group as defined earlier (AssING 2004c) may be paraphyletic and that the M. petrochilosi group should be included in the M. apicalis group.

From all the species of the M. apicalis group (including those of the former M. petrochilosi group), M. reliquus is readily distinguished by the much coarser and much more well-defined puncturation of the head and pronotum (somewhat resem- bling the condition in M. brunneus (Erichson)), by the chaetotaxy of the male stern- ite VII, as well as by the shape of the aedeagus.

From M. maronitus (widespread in the Eastern Mediterranean) and M. beydaghensis (southwestern Anatolia: W-Antalya, Isparta), the new species is addi- tionally separated by the uniformly reddish coloration, the absence of microsculp- ture on the forebody, the stouter modified setae at the posterior margin of the male sternite VII forming distinct patterns, as well as by the slightly different shape of the aedeagus.

The geographically closest representatives of the species previously attributed to the M. petrochilosi group - all of them with restricted distributions are M. caricus Fagel (southwestern and western Anatolia: Mugla, Izmir; Greece: Dhodhekänisos: Nikaria) and M. impar (Rhodos). From these species, M. reliquus is readily distin- guished by smaller body size, the absence of microsculpture on the forebody and consequently amuch more shiny appearance, the distinctly smaller eyes, the much shorter and more slender elytra, the less distinctly modified and less distinctly grouped setae at the posterior margin of the male sternite VII, as well as by the com- pletely different morphology of the aedeagus. For illustrations of the genitalia of the compared species see AssING (2004c).

Distribution and bionomics The species is apparently endemic to the Boz Daglar and the Aydin Daglari in western Anatolia (Fig. 71). Most of the specimens were sifted from the roots of grass and shrubs, some also from gravel and from oak leaf litter on grassy north slopes and in oak forests at altitudes of 850-1480 m. The type locality is illustrated in fig. 11 in ASSING (in press a).

ASSING, STAPHYLINIDAE FROM TURKEY V dey

Pseudomedon obscurellus (Erichson, 1840)

Material examined: Erzurum: 3 exs., 30-45km NNE Erzurum, Dumludagi, ca. 40°08N, 41°24E, 2200-2500 m, 14.V1.1998, leg. SOLODOVNIKOV (cAss, cSol).

This species is widespread in Europe (SMETANA 2004). It is here reported from Turkey for the first time.

Tetartopeus stylifer (Reitter, 1909)

Material examined: Erzurum: 1 ex., 35km NW Tortum, Mescit Daglari, ca. 40°30N, 41°25E, 1700-2000 m, 17.V1.1998, leg. SOLODOVNIKOV (cAss).

The species was previously known from Georgia, Ukraine, Russia, and Iraq (SMETANA 2004). It is here recorded from Turkey for the first time. The male geni- talia are illustrated by AssınG (2004b).

Xantholinus graecus Kraatz, 1858

Material examined: Antalya: 1 ex., Alanya env., 36°32N, 32°14E, 30.V.2004, leg. WEIGEL (cAss); 1 ex., Güzelbag, Zeytinbükü river, 36°40N, 31°53E, 160 m, 24.V.2004, leg. WEIGEL (cApf); 1 ex., 15km NE Alanya, Dimcayı river, 36°34N, 32°13E, 130m, 22.V.2004, leg. WEIGEL (cApf); 1 ex., Kemer near Antalya, 800 m, 1.V.1992, leg. BEHNE (DEI).

In Turkey, recent records of this species are known only from southern Anatolia (Mugla, Antalya, Mersin, Adana, Kahramanmaras, and Gaziantep provinces) (Ass- ING 2006f).

Xantholinus audrasi Coiffait, 1956

Material examined: Ankara: 4 exs., Gölbası, 750m, 29.X.1995, leg. Vrr (cAss). - Ordu: 1 ex., 25km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, road margin, roots of grass and herbs and moss sifted, 30.VII.2006, leg. AssınG (cAss).

In Turkey, this species was previously reported from Mugla, Antalya, Burdur, and Mersin provinces (AssING 2003a, 2006f).

Xantholinus osellai Bordoni, 1976 (Fig. 30)

Type material examined: Holotype d: Turchia leg. Osetia / Draganaz gec., VI.19[year illegible] / Xantholinus osellai mihi det. BorDont 1974 / Holotypus (MCSNV).

As is suggested by the similar highly derived internal structures of the aedeagus (presence of very long spine), X. osellai is the sister species of X. grandespinosus Ass- ing, from which it is distinguished as follows: body of smaller size; head smaller both absolutely and in relation to remainder of body (X. osellai: HW: 0.98 mm; HW/PW: 1.16; X. grandespinosus: HW: 1.15mm); eyes smaller, barely larger than antenno- mere I in cross-section (at widest point) (in X. grandespinosus distinctly larger); an- tennae shorter and with more transverse antennomeres, antennomeres VI-VII ap- proximately 3 times as wide as long (in X. grandespinosus about twice as wide as long); elytra without distinct microsculpture; aedeagus larger [despite smaller body size!] and with distinctly longer spine (length of spine: 1.04mm; in X. grandespinosus 0.53 mm) (Fig. 30).

16 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

Xantholinus grandespinosus Assing, 2006

Material examined: Ordu: 1 ex., 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, road margin, roots of grass and herbs and moss sifted, 30. VII.2006, leg. AssınG (cAss).

Previously, only the holotype (Amasya province) of this recently described species had become known (AssING 2006f).

Gabrius anatolicus Smetana, 1953

Type material examined: Holotype ¢: Yeniköy, Toros, 30.VIII.47, Anat. Exp. N. Mus. CSR. / Holotype / Gabrius Steph. anatolicus m., det. SMETANA 1952 / Mus. Nat. Pragae Inv. 18768 (NMP).

Additional material examined: 4 exs., Izmir, Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°07E, 1500 m, N-slope, Alnus and Salix litter sifted, 3.1V.2006, leg. AssınG (cAss).

Previously, only the type material of this species from southern Anatolia had be- come known; the above specimens represent the first record from western Anatolia.

Philonthus alpinus Eppelsheim, 1875

Material examined: Rize: 3 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). Artvin: 10 exs., ca. 40 km SW Artvin, source of Barhal river, ca. 40°05N, 41°30E, 2400m, 23.V1.1998, leg. SOLODOVNIKOV (cAss, cSol).

The species is here reported from Turkey for the first time.

Philonthus coprophilus Jarrige, 1949

Material examined: Erzurum: 4 exs., 30-45 km NNE Erzurum, Dumludagi, ca. 40°08N, 41°24E, 2200-2500 m, 14.V1.1998, leg. SOLODOVNIKOV (cAss, cSol).

The species was previously unknown from Turkey (HERMAN 2001).

Philonthus svanetiensis Coiffaıt, 1974

Material examined: Rize: 9 exs., 40km SSW Hopa, source of Caglayan D., ca. 41°06N, 41°22E, 2700-2900 m, 25.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 3 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 60km SSE Rize, Ovitdagi Gec., 40°38N, 40°45E, 2510 m, N- slope, under stones, 25.VII.2006, leg. AssınG (cAss); 1 ex., Ovitdagi Gec., 40°37N, 40°47E, 2710 m, N-slope, under stones, 25. V1I.2006, leg. Asstnc (cAss). Artvin: 1 ex., ca. 40 km SW Artvin, Barhal river valley, ca. 40°57N, 41°29E, 1800 m, 23.V1.1998, leg. SOLODOVNIKOV (cAss); 17 exs., ca. 40km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800 m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). Erzurum: 25 exs., 30-45 km NNE Erzu- rum, Dumludagi, ca. 40°11N, 41°27E, 2500-2900 m, 15.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex, 35km NW Tortum, Mescit Daglari, ca. 40°30N, 41°25E, 2100m, 18.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 35-40km NW Tortum, Mescit Daglari, ca. 40°30N, 41°17E, 2600 m, 19.V1.1998, leg. SOLODOVNIKOV (cAss).

According to SMETANA (2004), the species was previously unknown from Turkey.

ASSING, STAPHYLINIDAE FROM TURKEY V 17

Quedius (Raphirus) harpago n.sp. (Figs. 31-40, 50)

Types _ Holotype 6: TR Prov.: Izmir (12), NO Odemis, Boz Dag, 1560m, 21.IV.2006,

N38°20’11”, E28°6'26", leg. MEYBoHM & BracHat / Holotypus d Quedius harpago sp. n. det. V. AsstNG 2006 (cAss).

Paratype @: N38°20’10 E028°06'25 (12), Türkei, Aydin [recte: Izmir], Bozdag, 1300-1560 m, 21.IV.2006, 1. BRACHAT & MEYBOHM (cAss).

i) |

*

ne ie Fa Ba as am, = 4 TE g s an ee | " ve | N: A } f : T r La “A iJ

35 36 39

Figs. 30-40. Holotypes of Xantholinus osellai (30) and Quedius harpago n.sp. (31-39). - 30. Aedeagus. 31. Habitus. 32. Forebody. 33. Antenna. 34. Abdomen. 35-36. Median lobe of aedeagus in lateral and in ventral view. 37. Paramere. 38-39. Apical part of median lobe in ven-

tral and in lateral view. 40. Internal structures of aedeagus. Scale bars: 1mm (31-32, 34), 0.5 mm (30, 33, 35-37), 0.2 mm (38-40).

18 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700

Etymology The name (Latin, noun in apposition: grapnel, grappling hook) alludes to the conspicuous shape of the apex of the median lobe of the aedeagus.

Description

Measurements (inmm) and ratios (holotype, paratype): AL: 2.01, -; HL: 1.10, 1.13; HW: 1.14, 1.22; PW: 1.59, 1.62; PL: 1.46, 1.53; EL: 0.84, 0.82; EW: 1.62, 1.74; AW: 1.46, 1.65; TiL: 1.19, 1.25; TaL: 1.04, 1.07; ML: 1.40, -; TL: 8.9, 8.5; HL/HW: 0.96, 0.93; PW/HW: 1.39, 1.33; PW/PL: 1.08, 1.06; EL/PL: 0.57, 0.54; EW/PW: 1.02, 1.08; AW/EW: 0.91, 0.96; TiL/TaL: 1.15, 1.17.

Habitus as in Fig. 31. Coloration: body blackish; tarsi, pro- and mesofemora red- dish brown; protibia reddish brown, latero-ventrally weakly blackened; mesotibia reddish brown, latero-ventrally distinctly blackened; metafemur and metatibia blackish, with the base of the metatibia narrowly rufous; antennae reddish, with the apical half more or less infuscate, apical half of antennomere II weakly to distinctly darkened.

Head weakly transverse (Fig. 32); frons without punctures; integument with very fine microsculpture composed of isodiametric meshes anteriorly and of transverse striae posteriorly. Antenna as in Fig. 33, antennomere III slightly longer than II.

Pronotum distinctly wider than head and weakly transverse, with fine and shal- low transverse microsculpture, also near anterior angles.

Elytra approximately as wide as and at suture only little